User blog comment:Lord Loss/Monster Appreciation Day: Kushala Daora/@comment-4094047-20120220120018/@comment-969093-20120220153729

It's not a sub. Just a Daora whose skin has gone rusty and he has yet to shed it like he does in the Ecology video. They're not significantly more powerful (if at all) and their main elemental weakness is the Water Element as opposed to Dragon.